Thousands of staff at the Ministry in Longbenton have been striking against cuts which their union claims will undermine efforts to clamp down on tax avoidance.
PCS general secretary Mark Serwotka said: "It is sickening to see millionaires in the cabinet wringing their hands about the immorality of tax avoidance when it is their lack of political will to act that means we lose tens of billions of pounds every year.
"The case for investment in our public services as an alternative to austerity could not be more obvious than it is in HMRC.
"Yet the government wants to cut 10,000 more jobs from the department, letting the wealthy tax dodgers off the hook and punishing the rest of us for a recession we did not cause."
An HM Revenue & Customs spokesman said: "HMRC is disappointed with the decision to strike and will do everything it can to maintain services to the public. We are seeking dialogue with the PCS to address their concerns and will work to minimise any disruption to our customers.
"In our 2010 spending review the Government made £917m available to us to tackle avoidance, evasion and fraud.
"This is being used to increase our tax take from compliance work by £7bn a year in 2014/15 which we are on target to do.
"Last year alone we increased the yield from our compliance work to £13.9bn."
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